Department/Program Home Page


Department/Program Description

Neuroscience is a discipline that probes one of the last biological frontiers in understanding ourselves. It asks fundamental questions about how the brain and nervous system work in the expression of behavior. As such, the field takes on a clear interdisciplinary character: All scientific levels of organization (behavioral, developmental, molecular, cellular, and systems) contribute to our understanding of the nervous system. Neuroscience has been a field of particularly active growth and progress for the past two decades, and it is certain to be an area where important and exciting developments will continue to occur. At Wesleyan, the neurosciences are represented by the teaching and research activities of faculty members in the departments of biology, psychology, and chemistry. The neuroscience and behavior (NS&B) curriculum is both comprehensive and provides diverse approaches to learning. Through lecture/seminars, lab-based methods courses, and hands-on research experience, students are afforded a rich educational experience. Unique among schools of comparative size, Wesleyan has small but active graduate programs leading to BA/MA and PhD degrees. This attribute, together with the high success rate of faculty in obtaining research grant support, further enhances the education of undergraduates by providing additional mentoring, more research opportunities, and access to state-of-the-art laboratories. The mission of the NS&B program is to provide the foundation for a variety of career options in science, medicine, and private industry. For more information, see

Admission to the Major

One or more of the foundation courses in biology (BIOL181, BIOL182) are prerequisites for the advanced NS&B courses offered by the Biology Department. Although not legislated as prerequisites, NS&B213 and NS&B laboratory courses provide important conceptual and practical background for independent research in the junior and senior years. The ideal course sequence would include BIOL181 and BIOL182 along with chemistry in the first year. In the sophomore year, one would take NS&B213. The other required courses and research tutorials would be spread out over the last two years. For information on the pathway through the major, please visit for further information.

To be admitted to the major during March of the sophomore year, a student must have completed, with grades of C- or better, at least two of the full-credit courses listed in foundation and core courses that follow. At least one of these credits must be either NS&B213 or BIOL181.

Major Requirements




Five advanced courses from the following list are required for students; two must be cross-listed with biology; two cross-listed with psychology; and one, a research tutorial or methodological course. Some courses appear in both Biology and Psychology lists but may be counted only once, in either category.   

Cross-listed with biology

  • NS&B224 Hormones, Brain, and Behavior
  • NS&B239 Functional Anatomy of the Human Brain*
  • NS&B245 Cellular Neurophysiology
  • NS&B252 Cell Biology of the Neuron
  • NS&B254 Comparative Animal Behavior
  • NS&B299 Waves, Brains, and Music
  • NS&B303 Receptors, Channels, and Pumps: Advanced Topics in Membrane Protein Structure and Function
  • NS&B317 Neuroethics
  • CHEM323/NS&B323 Biochemistry of Neurodegenerative Disease 
  • NS&B325 Stem Cells: Basic Biology to Clinical Application
  • NS&B328 Chemical Senses
  • NS&B343 Muscle and Nerve Development
  • NS&B345 Developmental Neurobiology
  • NS&B347 Mammalian Cortical Circuits
  • NS&B351 Neurobiology of Learning and Memory
  • NS&B357 Sex and Gender: From Synapse to Society
  • NS&B353 Neurobiology of Neurological Disorders*
  • NS&B356  Neurodevelopmental Disorders*
  • NS&B360 Calderwood Seminar in Public Writing: Neuroplasticity and the Brain

Cross-listed with psychology

  • NS&B220 Cognitive Psychology
  • NS&B221 Human Memory
  • NS&B222 Sensation and Perception
  • NS&B225 Cognitive Neuroscience
  • NS&B227 Motivation and Reward
  • NS&B228 Clinical Neuropsychology
  • NS&B239 Functional Anatomy of the Human Brain*
  • NS&B316 Schizophrenia and Its Treatment: Neuroscientific, Historical, and Phenomenological Perspectives
  • NS&B317 Neuroethics
  • NS&B329 Neural Costs of War
  • NS&B341 Psychology of Learning and Memory
  • NS&B342 Music Perception and Cognition
  • NS&B348 Origins of Knowledge
  • NS&B353 Neurobiology of Neurological Disorders*
  • NS&B356 Neurodevelopmental Disorders*

Research methods and practica

  • BIOL242 Quantitative Methods for the Biological and Environmental Sciences
  • MATH132 Elementary Statistics
  • NS&B210 Research Methods in Cognition
  • NS&B215 Research Methods: Behavioral Methods in Animal Research
  • NS&B243 Neurohistology
  • NS&B250 Laboratory in Cellular and Behavioral Neurobiology
  • NS&B280 Applied Data Analysis
  • NS&B383 Advanced Research in Learning and Memory
  • NS&B390 Experimental Investigations into Reading
  • NS&B392 Behavioral Methods in Affective Neuroscience
  • NS&B398 Advanced Research in Auditory Cognitive Neuroscience
  • NS&B399 Lab in Gambling, Drugs, and Junk Food
  • NS&B409/NS&B410 Senior Thesis Tutorial or NS&B423/NS&B424 Advanced Research Seminar for two semesters, both in the lab of the same faculty member
  • PSYC200 Statistics: An Activity-Based Approach

Note: MATH132 can be taken to meet requirements for either the methodological or foundation major requirements, but not both.  Methodological courses cannot be credited toward the requirements of advanced courses cross-listed with biology or psychology. *Courses listed in both categories A. or B. can be counted only in A. or B. but not both

Courses of relevance outside the program. Though not requirements of the major, students should be aware that courses in organic chemistry and molecular biology, as well as courses in non-neuroscience areas of biology and psychology, complement the NS&B major and should be considered, in consultation with your advisor, when planning your program of study. 


Foundation courses: A student who has taken foundation courses outside of Wesleyan may be able to apply them to the major. As a general rule, courses acceptable to the biology, chemistry, and physics departments for university credit are acceptable to the NS&B program for substitution for foundation courses.

Advanced courses: Advanced courses, inside or outside of the University, might be acceptable as substitutes for the advanced courses of the NS&B major. In general, only one such course can be substituted, and approval must be obtained in advance from the program director.


NS&B majors are encouraged to become involved in the research of the faculty. Research tutorials and senior thesis tutorials are taken with mode of grading and amount of credit to be arranged with the research supervisor. Research tutorials are numbered NS&B411/NS&B412, NS&B409/NS&B410, and NS&B423/NS&B424. These courses can fulfill the research methods requirement or can receive graduation credit.  For the most up-to-date information on NS&B faculty research, please visit our department website.

Student Learning Goals

Our program offers a curriculum that encourages fluency across multiple disciplines in the field of neuroscience and behavior. Immersion in this field requires thinking across multiple levels of analysis and an appreciation for how complex and broad questions can be made amenable to scientific inquiry. In terms of goals, we have three areas of knowledge that we expect all students to acquire by the time they have completed the NS&B major:

  • Structure: The parts and how they connect. Structural knowledge includes neural development, neuroanatomy, neurotransmitters, and the cell and molecular biology of the neuron.
  • Function: How the parts come together to produce systems. Such systems include various sensory, motor, and neuroendocrine systems. Knowledge concerning function is gained by studies of structures and studies of perception, learning and memory, behavior, and cognition.
  • Theory: Governing principles that can be proposed from all the above. Examples of theories include those that address the relationships between brain and behavior, articulate how brain structure and function changes over time, and explain cognitive and perceptual processes.

In addition, it is our goal that all students can skillfully apply and analyze knowledge gained from their studies. Statistics courses, lab-based methods courses, and/or direct experience in research projects serve this goal.

Advanced Placement
AP credit may be used to place out of any of the foundation courses, subject to the guidelines of the department hosting these courses.
George H. Acheson and Grass Foundation Prize in Neuroscience: Established in 1992 by a gift from the Grass Foundation, this prize is awarded to an outstanding undergraduate in the Neuroscience and Behavior Program who demonstrates excellence in the program and who also shows promise for future contributions in the field of neuroscience.
BA/MA Program
This program provides an attractive option for science majors to enrich their course and research background. Students are advised to begin research by their junior year if they intend to pursue the BA/MA. Admission is competitive and based on GPA, faculty recommendations, and research experience. For more information, please visit
Additional Information
  • Teaching apprenticeships. Students may be appointed teaching apprentices with the approval of the participating faculty member and the Office of Academic Affairs. The apprenticeship position involves assisting a faculty member in the teaching of a course. Concurrently, the apprentice enrolls in an apprenticeship tutorial (NS&B491/NS&B492) that is usually a one-credit course and operates in either the graded or credit/no credit mode.
  • Petitioning for exemptions. A student may request a variance from the requirements of the major or for honors by submitting a written petition to the chair of the program. The petition should indicate why the requirement cannot be met and the educational justification for the alternative. The petition will be considered by the NS&B faculty, and the student will receive a statement of the decision by letter.
  • Seminars. The program periodically invites neuroscientists from outside Wesleyan to come here and describe their research. These seminars frequently complement course material and give students the opportunity to interact with noted researchers. The talks are usually scheduled for noon on Wednesdays. Students are encouraged to attend.

To be considered for honors, a student must be an NS&B major and have a B average (grade average 85) in the courses credited to the major. The student must submit a laboratory research thesis that was supervised by a member of the NS&B faculty and be recommended for honors by the NS&B faculty.

General Introduction


The Neuroscience & Behavior Program offers graduate work leading to the degree of Master of Arts through the BA/MA program.  The program has a strong research orientation.  It also includes course work, seminars, and, in some cases, teaching.  A student hoping to enter this program will be expected to declare the intention to do so in the first semester of their junior year to permit the design of an acceptable program with both the major department and a research advisor within that department. 



The MA will require a minimum of 6 credits in addition to the 32 necessary for the Wesleyan BA. Three credits will be earned through Journal Club I & II (0.25 credits x 2 = 0.50), Advanced Research NS&B549/NS&B550 (1.0 credits x 2 = 2.0) and BIOL557 (.050 credits).  The remaining credits will be in 1-credit lab, lecture or seminar courses (200, 300, and 500) as determined by the student and faculty advisor.  MA credit will only be awarded for academic work in which grades of B- or higher have been earned. A student in the program who earns more than 32 credits in four years may apply any excess credits toward the MA, providing that they are in the major area and they have not been used to fulfill the undergraduate major requirement. 

Progress and Qualifying Exams
A 3-member committee of the faculty will be established upon acceptance in the BA/MA program. The candidate will meet with their committee in early stages of research and meet with them in the second semester of their MA year. This committee determines when sufficient experimental work has been completed and must approve the final written document. Students in this program are required to submit a MA thesis describing the research which they have carried out in partial fulfillment of the degree requirements. 
There are no requirements for BA/MA candidates to teach although the opportunity may arise.   
Students in this program will submit an MA thesis describing the research and will present this to their committee in a closed-door oral defense of the thesis.  Following the oral defense with the thesis committee, the students will receive two grades for their thesis work: one for the oral component and one for the written component of the thesis. Additionally, students are required give a public presentation during the BIOL557 seminar describing their research as partial fulfillment of the degree requirements.