About the Major

Department/Program Description

These are thrilling times to study biology. Advances in molecular biology, epigenetics, and bioinformatics are leading to extraordinary new insights in every field, from evolution and ecology to development, cell biology, genetics/genomics, and neuroscience. These research areas are providing essential information as we address the urgent challenges of biodiversity conservation, global climate change, epidemiology, and human health and well-being. Biology is also at the heart of new ways of understanding ourselves as human beings in relation to other living things. Connections between biological disciplines are raising key questions in new ways, while biological knowledge has become fundamentally integrated with social and medical ethics, public policy, and journalism.

The Biology Department offers a broad range of courses that emphasize the process of scientific inquiry and current experimental approaches. Our courses also consider real-world implications of biological issues: the ethics of embryonic stem cell research, gender issues and reproductive technologies, the AIDS epidemic, and the impact of human activity on natural communities. Biology courses can be the start of a dedicated career in research, medicine, conservation, public health, bioethics, sustainable resource use, and many other areas. They can also bring the intellectual excitement of these investigations to students whose major focus is in the arts, humanities, or social sciences. We welcome students of all backgrounds and interests to join us.

Courses for Non-Majors

The following courses do not have prerequisites and, as such, are appropriate for non-majors.

BIOL106 The Biology of Sex 1
BIOL137 Writing About Evolution 1
BIOL140 Classic Studies in Animal Behavior 1
BIOL145F Primate Behavior: The Real Monkey Business (FYS) 1
BIOL149 Neuroethology: Sensory Basis of Animal Orientation and Navigation 1
BIOL173 Global Change and Infectious Disease 1
BIOL181 Principles of Biology I: Cell Biology and Molecular Basis of Heredity 1
BIOL182 Principles of Biology II 1
BIOL186 Introduction to the Biology of Nutrition and Impact on Human Health 1
BIOL197 Introduction to Environmental Studies 1
Student Learning Goals
The Biology Department expects its majors to develop a broad and integrative understanding of the theory and practice of biology across a range of disciplines and levels of biological organization. The curricular requirements of the major are designed to provide enough flexibility for each student to choose a disciplinary emphasis of most interest and fulfill the additional expectation of achieving some depth of knowledge in a particular area through a relatively intensive classroom or laboratory experience. In this context, we want our students to develop skills in critical and quantitative thinking, creative problem-solving, and intuition for the process of scientific reasoning. We also encourage our students to engage in ethical thinking about biological research and the role of biology in society and sustainability. A complete program of study in biology entails the application of these skills to designing or conducting original research (including scholarly research via scientific databases), writing about and orally communicating scientific concepts, as well as the comprehension and critical interpretation of primary scientific literature. Our ultimate goal is, therefore, to train students to use their biological knowledge and skills to become effective, scientifically informed citizens and professionals.
Admission to the Major

Students are encouraged to begin their major in the first year so that they can take maximum advantage of upper-level biology courses and research opportunities in later years. However, the major can certainly be successfully completed if begun during sophomore year, and many students are able to combine the biology major with a semester abroad.

A prospective biology major begins with a series of two core introductory courses. Students should begin the core series with BIOL181 and its associated laboratory course, BIOL191, which are offered in the fall semester. BIOL181 is offered in a number of small sections rather than a single large lecture class. These small sections allow for problem-based learning at a more individualized pace as students master the first semester of university-level biology. Students should enroll separately for the lab course, BIOL191. These courses do not have prerequisites or corequisites, but it is useful to have some chemistry background or to take chemistry concurrently. In the spring semester, the prospective major should take BIOL182and its laboratory course, BIOL192. An optional spring course, BIOL194, is offered to students of BIOL182 who wish a challenging reading and discussion experience in addition to the lectures.

Major Requirements

The biology major's program of study consists of the following:

Note: No more than three of these mid-level courses (listed above) may be counted towards the six advanced elective requirement.

  • IMPORTANT INFORMATION FOR DOUBLE MAJORS: At least two elective courses (200-level and above) that are counted toward the biology major cannot be simultaneously used to fulfill any other major.
  • In addition, if a student is double-majoring in biology and NS&B, NS&B/BIOL213 cannot count toward the six electives required for the biology major; however, NS&B/BIOL213 will fulfill the Column 2 breadth requirement.
  • Two semesters of general chemistry (CHEM141-CHEM142 or CHEM143-CHEM144)
  • Any three additional semesters of related courses from at least two different departments:

  • Biology majors are allowed to apply at most one elective course or one cognate course taken credit/unsatisfactory toward fulfilling the major requirements; however, this is discouraged because good performance in major courses is an important aspect of a student's transcript.
  • A strong chemistry background is especially recommended for students planning to enter graduate or medical school. Most medical and other health-related graduate schools require two years of college-level chemistry, including laboratory components, as well as a course in biochemistry.
  • Students planning to go on to medical, dental, or other health professions graduate school should note that a year each of introductory biology, physics, and math (such as calculus or statistics) and two years of chemistry (general and organic) are required for admission, including any laboratory components.

Electives may be chosen from among the following courses at the 200, 300, or 500 levels. See WesMaps for current course offerings. The courses are grouped thematically for your convenience only.


BIOL212 Principles and Mechanisms of Cell Biology 1
BIOL218 Developmental Biology 1
MB&B/BIOL232 Immunology 1
MB&B/BIOL237 Signal Transduction 1
BIOL245 Cellular Neurophysiology 1
BIOL334 Shaping the Organism 1
BIOL325 Stem Cells: Basic Biology to Clinical Application 1
BIOL340/540 Issues in Development and Evolution 1
BIOL343/543 Muscle and Nerve Development 1
BIOL/NS&B345 Developmental Neurobiology 1


BIOL214 MacroEvolution, Pattern and Process 1
BIOL215 Evolution in Human-Altered Environments 1
BIOL216 Ecology 1
BIOL220 Conservation Biology 1
BIOL235 Comparative Vertebrate Anatomy 1
BIOL254 Comparative Animal Behavior 1
BIOL290 Plant Form and Diversity 1
BIOL310 Genomics Analysis 1
BIOL316/516 Plant-Animal Interactions 1
BIOL318/518 Nature and Nurture: The Interplay of Genes and Environment 1
BIOL327/527 Evolutionary and Ecological Bioinformatics 1
BIOL340/540 Issues in Development and Evolution 1
BIOL346 The Forest Ecosystem 1


MB&B208 Molecular Biology 1
BIOL210 Genomics: Modern Genetics, Bioinformatics, and the Human Genome Project 1
BIOL/MB&B265/COMP113/CIS265 Bioinformatics Programming 1
BIOL327/527 Evolutionary and Ecological Bioinformatics 1
MB&B/BIOL231 Microbiology 1
BIOL310 Genomics Analysis 1
BIOL327/527 Evolutionary and Ecological Bioinformatics 1
MB&B333/533 Gene Regulation 1
MB&B394 Advanced Laboratory in Molecular Biology and Genetics 1
E&ES280 Introduction to GIS 1
MB&B306 Epigenetics 1


NS&B/BIOL213 Behavioral Neurobiology 1
BIOL/NS&B224 Hormones, Brain, and Behavior 1
BIOL235 Comparative Vertebrate Anatomy 1
BIOL/NS&B239 Functional Anatomy of the Human Brain 1
NS&B/BIOL243 Neurohistology 1
BIOL/NS&B245 Cellular Neurophysiology 1
BIOL/NS&B247 Laboratory in Neurophysiology 1
BIOL/NS&B250 Laboratory in Cellular and Behavioral Neurobiology 1
BIOL/NS&B252 Cell Biology of the Neuron 1
NS&B/BIOL254 Comparative Animal Behavior 1
BIOL290 Plant Form and Diversity 1
BIOL/NS&B299 Waves, Brains, and Music 1
NS&B/BIOL328 Chemical Senses 1
BIOL/NS&B345 Developmental Neurobiology 1
BIOL/NS&B351 Neurobiology of Learning and Memory 1
NS&B/BIOL353 Neurobiology of Neurological Disorders 1
NS&B/BIOL360 Calderwood Seminar in Public Writing: Neuroplasticity and the Brain 1


Cross-listed courses that are included on the list above are automatically credited to the biology major. At least two elective courses (200-level and above) that are counted toward the biology major must be used to fulfill only the biology major and cannot be simultaneously used to fulfill another major.

Depending on the student’s specific program, and with prior permission of the chair, up to two biology courses from outside the department may be counted toward the major. Two Wesleyan courses that fall into this category are ANTH202 and ANTH349.

Additional courses that may be credited to the major: BIOL242, E&ES234, MB&B228, CHEM323 and CHEM383, and ECON300. MB&B228 may be counted as long as neither MB&B208 nor CHEM383 is counted toward the major.

Biology majors are allowed to apply at most one elective course taken credit/unsatisfactory toward fulfilling the major requirements; however, this is discouraged because good performance in major courses is an important aspect of a student’s transcript.

Courses in the BIOL 400 series (such as research tutorials) contribute toward graduation but do not count toward the major.

Capstone Experience
As a capstone experience, senior biology majors are encouraged to participate in one of the following opportunities for intensive scientific engagement: a hands-on laboratory or field course, participation in lab research (typically begun prior to senior year), or enrollment in an advanced (300-level) seminar or class. A series of faculty-student dinners during fall and spring of senior year provide further opportunities to discuss emergent scientific issues and approaches and their relation to students’ career goals.

To be considered for departmental honors, a student must

  • Be a biology major and be recommended to the department by a faculty member. It is expected that the student will have at least a B average (grade point average 85) in courses credited to the major.
  • Submit a thesis based on laboratory research, computational research, or mathematical modeling. The thesis is carried out under the supervision of a faculty member of the department.
Advanced Placement

Students who have received a grade of 4 or 5 on the AP exam may receive one University credit toward graduation.

Students with a score of 4 or 5 may place out of one of the two Introductory Biology courses - BIOL181 or BIOL182 but must first consult with an instructor teaching these courses. 

Students interested in placing out of MB&B181 in the fall semester should contact Professor Michelle Murolo (mmurolo@wesleyan.edu) regarding the placement exam. 

No follow-up course is required. These courses are considered essential background for our upper-level courses; students are highly encouraged to enroll in both semesters.

However, we recommend against this for almost all students, especially those who may be interested in the biology major. Although some of the MB&B181/BIOL181 material will be familiar from a high school AP course, the depth and rigor of MB&B181/BIOL181 provide a strong foundation as you move forward to more advanced courses. Alternatively, students with AP 4 or 5 may consult individually with the BIOL182 faculty regarding placing out of this second-semester introductory course. However, both courses are considered essential background for our upper-level courses; students are highly encouraged to enroll in both semesters.


Dr. Neil Clendeninn Prize. Established in 1991 by George Thornton, Class of 1991, and David Derryck, Class of 1993, for the African American student who has achieved academic excellence in biology and/or molecular biology and biochemistry. This student must have completed his or her sophomore year and in that time have exemplified those qualities of character, leadership, and concern for the Wesleyan community as shown by Dr. Neil Clendeninn, Class of 1971.

The Peirce Prize. Awarded in successive years for excellence in biology, chemistry, and geology.

Transfer Credit
Up to two outside credits for biology courses may also be applied from another institution (during a study abroad program, for example). Prior permission must be obtained from the departmental liaison Professor Michael Singer (msinger@wesleyan.edu) to ensure creditability of specific courses from other institutions. Information and a downloadable form can be found here https://www.wesleyan.edu/studentaffairs/pdfs/ptcapril2016.pdf
Related Programs or Certificates

Environmental Studies Certificate. The Environmental Studies (ENVS) program is interdisciplinary and offers both a certificate and a linked major. The ENVS linked major is a secondary major and requires a student to also have a primary major in another department, program, or college. ENVS majors write a senior thesis or essay in environmental studies that is mentored by a professor in another department, program, or college (e.g., biology). There is also an opportunity to earn an ENVS certificate, which does not require a senior thesis or essay. See: wesleyan.edu/coe/academics/.

Informatics and Modeling Certificate. The Integrative Genomic Science pathway within this certificate will be of particular interest for life science majors. See wesleyan.edu/imcp/igs.html.

Neuroscience and Behavior Program. Several faculty members in the Biology and Psychology Departments also participate in the Neuroscience and Behavior Program that, at the undergraduate level, constitutes a separate major. Information about that program can be found at wesleyan.edu/nsb.

The graduate program is an integral part of the Biology Department’s offerings. Not only are graduate students active participants in the undergraduate courses, but, also, upper-level undergraduates are encouraged to take graduate-level courses and seminars (500 series). Research opportunities are also available for undergraduates, and, frequently, these involve close interaction with graduate students.

Additional Information
The seminar series features distinguished scientists from other institutions who present lectures on their research findings. One objective of these seminars is to relate material studied in courses, tutorials, and research to current scientific activity. These seminars are usually held on Thursdays at noon and are open to all members of the University community. Undergraduates are especially welcome.